:root{
    --ink:#0B1614;
    --ink-2:#11201D;
    --ink-3:#1A2E29;
    --paper:#EDE6D6;
    --paper-dim:#A9A196;
    --bio:#5FE1B0;
    --bio-dim:#3B9E7C;
    --amber:#E0A64A;
    --rust:#C1573A;
    --line: rgba(237,230,214,0.14);
    --line-strong: rgba(237,230,214,0.28);
    --maxw: 1180px;
  }

  *{box-sizing:border-box;}
  html{scroll-behavior:smooth; scroll-padding-top:88px;}
  body{
    margin:0;
    background:
      radial-gradient(ellipse 900px 500px at 15% -10%, rgba(95,225,176,0.08), transparent 60%),
      radial-gradient(ellipse 700px 500px at 100% 10%, rgba(224,166,74,0.06), transparent 60%),
      var(--ink);
    color:var(--paper);
    font-family:'Inter', sans-serif;
    font-size:16px;
    line-height:1.6;
    -webkit-font-smoothing:antialiased;
  }
  body::before{
    content:"";
    position:fixed; inset:0;
    background-image:
      linear-gradient(var(--line) 1px, transparent 1px),
      linear-gradient(90deg, var(--line) 1px, transparent 1px);
    background-size:64px 64px;
    opacity:0.25;
    pointer-events:none;
    z-index:0;
    mask-image: radial-gradient(ellipse 80% 60% at 50% 0%, black 20%, transparent 75%);
  }
  a{color:inherit;}
  img,svg{max-width:100%; display:block;}
  ::selection{ background:var(--bio); color:var(--ink); }
  :focus-visible{ outline:2px solid var(--bio); outline-offset:3px; }

  h1,h2,h3{ font-family:'Space Grotesk', sans-serif; margin:0; font-weight:600; letter-spacing:-0.01em; }
  .taxon{ font-family:'Source Serif 4', serif; font-style:italic; color:var(--bio); }
  .mono{ font-family:'IBM Plex Mono', monospace; }
  .eyebrow{
    font-family:'IBM Plex Mono', monospace;
    text-transform:uppercase;
    letter-spacing:0.14em;
    font-size:0.72rem;
    color:var(--amber);
    display:flex; align-items:center; gap:10px;
    margin-bottom:18px;
  }
  .eyebrow::before{
    content:""; width:7px; height:7px; border-radius:50%;
    background:var(--amber); flex:none;
  }

  .wrap{ max-width:var(--maxw); margin:0 auto; padding:0 28px; position:relative; z-index:1; }
  section{ padding:120px 0; position:relative; z-index:1; border-bottom:1px solid var(--line); }
  section:last-of-type{ border-bottom:none; }

  /* NAV */
  .nav{
    position:fixed; top:0; left:0; right:0; z-index:50;
    background:rgba(11,22,20,0.72);
    backdrop-filter:blur(10px);
    border-bottom:1px solid var(--line);
  }
  .nav .wrap{ display:flex; align-items:center; justify-content:space-between; padding-top:18px; padding-bottom:18px; }
  .logo{ font-family:'IBM Plex Mono', monospace; font-weight:500; letter-spacing:0.02em; font-size:1rem; display:flex; align-items:center; gap:9px; text-decoration:none; }
  .logo .dot{ width:8px; height:8px; border-radius:50%; background:var(--bio); box-shadow:0 0 0 0 rgba(95,225,176,0.6); animation:pulse-dot 2.6s ease-out infinite; }
  @keyframes pulse-dot{
    0%{ box-shadow:0 0 0 0 rgba(95,225,176,0.55); }
    70%{ box-shadow:0 0 0 8px rgba(95,225,176,0); }
    100%{ box-shadow:0 0 0 0 rgba(95,225,176,0); }
  }
  .navlinks{ display:flex; gap:34px; list-style:none; margin:0; padding:0; font-size:0.9rem; }
  .navlinks a{ text-decoration:none; color:var(--paper-dim); transition:color .2s; }
  .navlinks a:hover{ color:var(--bio); }
  .navbtn{ display:none; background:none; border:1px solid var(--line-strong); color:var(--paper); width:40px; height:40px; border-radius:6px; cursor:pointer; }

  .btn{
    font-family:'IBM Plex Mono', monospace;
    font-size:0.82rem;
    text-decoration:none;
    display:inline-flex; align-items:center; gap:8px;
    padding:13px 22px;
    border-radius:3px;
    border:1px solid var(--line-strong);
    color:var(--paper);
    transition:border-color .2s, background .2s, color .2s;
    white-space:nowrap;
  }
  .btn:hover{ border-color:var(--bio); color:var(--bio); }
  .btn.solid{ background:var(--bio); color:var(--ink); border-color:var(--bio); font-weight:500; }
  .btn.solid:hover{ background:var(--paper); border-color:var(--paper); color:var(--ink); }

  /* HERO */
  .hero{ padding-top:190px; padding-bottom:100px; }
  .hero-grid{ display:grid; grid-template-columns:1.1fr 0.9fr; gap:70px; align-items:center; }
  .hero h1{ font-size:clamp(2.1rem, 4.4vw, 3.4rem); line-height:1.12; margin-bottom:26px; }
  .hero h1 em{ font-style:normal; color:var(--bio); }
  .hero p.lead{ font-size:1.08rem; color:var(--paper-dim); max-width:52ch; margin-bottom:36px; }
  .hero-ctas{ display:flex; gap:14px; flex-wrap:wrap; margin-bottom:46px; }
  .specimen-tag{
    font-family:'IBM Plex Mono', monospace;
    font-size:0.76rem;
    color:var(--paper-dim);
    border:1px dashed var(--line-strong);
    padding:14px 18px;
    display:inline-block;
    max-width:100%;
  }
  .specimen-tag b{ color:var(--bio); font-weight:500; }
  .specimen-tag .sep{ color:var(--line-strong); margin:0 6px; }

  .hero-visual{
    aspect-ratio:1/1;
    border:1px solid var(--line);
    border-radius:6px;
    position:relative;
    overflow:hidden;
    background:linear-gradient(160deg, var(--ink-2), var(--ink));
  }
  .hero-visual .ring{
    position:absolute; border-radius:50%; border:1px solid var(--line-strong);
  }
  .hero-visual .core{
    position:absolute; top:50%; left:50%; width:14px; height:14px; margin:-7px;
    border-radius:50%; background:var(--bio);
    box-shadow:0 0 40px 6px rgba(95,225,176,0.5);
    animation:core-pulse 3.2s ease-in-out infinite;
  }
  @keyframes core-pulse{ 0%,100%{ transform:scale(1); } 50%{ transform:scale(1.35); } }
  .hero-visual .label{
    position:absolute; font-family:'IBM Plex Mono', monospace; font-size:0.68rem; color:var(--paper-dim);
    letter-spacing:0.06em;
  }

  /* SECTION HEADS */
  .head{ max-width:640px; margin-bottom:56px; }
  .head h2{ font-size:clamp(1.6rem,3vw,2.2rem); margin-bottom:16px; }
  .head p{ color:var(--paper-dim); font-size:1rem; margin:0; }

  /* LINEAGE / CLADOGRAM */
  .tree-frame{
    border:1px solid var(--line);
    border-radius:6px;
    padding:20px 10px;
    overflow-x:auto;
    background:var(--ink-2);
    margin-bottom:52px;
  }
  .tree-frame svg{ min-width:980px; width:100%; height:auto; display:block; margin:0 auto; }
  .branch-line{ fill:none; stroke:var(--line-strong); stroke-width:1.6; }
  .branch-line.active{ stroke:var(--bio); }
  .node-circle{ fill:var(--ink-2); stroke:var(--paper-dim); stroke-width:1.6; }
  .node-circle.active{ stroke:var(--bio); fill:var(--ink); }
  .node-label{ font-family:'Space Grotesk', sans-serif; font-size:15px; fill:var(--paper); font-weight:500; }
  .node-sub{ font-family:'Source Serif 4', serif; font-style:italic; font-size:12px; fill:var(--paper-dim); }
  .node-now{ font-family:'IBM Plex Mono', monospace; font-size:11px; fill:var(--amber); letter-spacing:0.06em; }

  .draw{
    stroke-dasharray:900;
    stroke-dashoffset:900;
    animation:draw 1.6s ease forwards;
  }
  @keyframes draw{ to{ stroke-dashoffset:0; } }
  .pop{ opacity:0; transform:scale(0.6); transform-origin:center; animation:pop .5s ease forwards; }
  @keyframes pop{ to{ opacity:1; transform:scale(1); } }

  .lineage-grid{ display:grid; grid-template-columns:repeat(3,1fr); gap:1px; background:var(--line); border:1px solid var(--line); border-radius:6px; overflow:hidden; }
  .lineage-card{ background:var(--ink-2); padding:26px 24px; }
  .lineage-card .num{ font-family:'IBM Plex Mono', monospace; font-size:0.72rem; color:var(--amber); margin-bottom:10px; display:block; }
  .lineage-card h3{ font-size:1.02rem; margin-bottom:6px; }
  .lineage-card .taxon{ font-size:0.86rem; display:block; margin-bottom:12px; }
  .lineage-card p{ color:var(--paper-dim); font-size:0.88rem; margin:0; }
  .lineage-card.now{ background:var(--ink-3); }
  .lineage-card.now .num{ color:var(--bio); }

  /* SERVICES */
  .cards{ display:grid; grid-template-columns:repeat(2,1fr); gap:1px; background:var(--line); border:1px solid var(--line); border-radius:6px; overflow:hidden; }
  .card{ background:var(--ink-2); padding:36px; transition:background .25s; }
  .card:hover{ background:var(--ink-3); }
  .card .tag{ font-family:'IBM Plex Mono', monospace; font-size:0.7rem; color:var(--amber); letter-spacing:0.08em; margin-bottom:16px; display:block; }
  .card h3{ font-size:1.15rem; margin-bottom:12px; }
  .card p{ color:var(--paper-dim); font-size:0.92rem; margin:0; }

  /* PROJECTS */
  .projects-list{ display:flex; flex-direction:column; border-top:1px solid var(--line); }
  .project-row{
    display:grid; grid-template-columns:100px 1fr auto; gap:28px; align-items:start;
    padding:34px 0; border-bottom:1px solid var(--line);
  }
  a.project-row{ transition:background .2s; }
  a.project-row:hover{ background:var(--ink-2); }
  a.project-row:hover h3{ color:var(--bio); }
  .project-row .code{ font-family:'IBM Plex Mono', monospace; font-size:0.78rem; color:var(--paper-dim); padding-top:4px; }
  .project-row h3{ font-size:1.15rem; margin-bottom:8px; }
  .project-row .taxon{ display:block; font-size:0.9rem; margin-bottom:10px; }
  .project-row p{ margin:0; color:var(--paper-dim); font-size:0.92rem; max-width:60ch; }
  .status{
    font-family:'IBM Plex Mono', monospace; font-size:0.68rem; letter-spacing:0.06em;
    color:var(--bio); border:1px solid var(--bio-dim); padding:6px 10px; border-radius:20px;
    white-space:nowrap; height:fit-content;
  }

  /* CONTACT */
  .contact-grid{ display:grid; grid-template-columns:1.2fr 0.8fr; gap:60px; align-items:end; }
  .contact h2{ font-size:clamp(1.8rem,3.6vw,2.6rem); margin-bottom:20px; }
  .contact p.lead{ color:var(--paper-dim); max-width:48ch; margin-bottom:32px; }
  .contact-block{ border:1px solid var(--line); border-radius:6px; padding:28px; }
  .contact-block .row{ display:flex; justify-content:space-between; padding:12px 0; border-bottom:1px solid var(--line); font-size:0.92rem; }
  .contact-block .row:last-child{ border-bottom:none; }
  .contact-block .row span:first-child{ color:var(--paper-dim); font-family:'IBM Plex Mono', monospace; font-size:0.76rem; text-transform:uppercase; letter-spacing:0.06em; }
  .contact-block .row a{ text-decoration:none; color:var(--bio); }

  footer{ padding:40px 0; }
  footer .wrap{ display:flex; justify-content:space-between; align-items:center; flex-wrap:wrap; gap:12px; }
  footer .mono{ font-size:0.76rem; color:var(--paper-dim); }

  .reveal{ opacity:0; transform:translateY(18px); transition:opacity .7s ease, transform .7s ease; }
  .reveal.in{ opacity:1; transform:translateY(0); }

  @media (max-width:880px){
    .hero-grid{ grid-template-columns:1fr; }
    .hero-visual{ max-width:340px; margin:0 auto; }
    .contact-grid{ grid-template-columns:1fr; }
    .cards{ grid-template-columns:1fr; }
    .lineage-grid{ grid-template-columns:1fr; }
    .navlinks{ display:none; }
    .navbtn{ display:block; }
    .project-row{ grid-template-columns:1fr; }
    .status{ justify-self:start; }
    section{ padding:80px 0; }
  }
  .navlinks.open{ display:flex; position:absolute; top:64px; left:0; right:0; background:var(--ink-2); flex-direction:column; padding:20px 28px; gap:18px; border-bottom:1px solid var(--line); }

  @media (prefers-reduced-motion: reduce){
    *{ animation-duration:0.001ms !important; animation-iteration-count:1 !important; transition-duration:0.001ms !important; }
    .draw{ stroke-dashoffset:0; }
  }

  /* NAV disabled item (current page indicator) */
  .navlinks .navlink-disabled{
    color:var(--paper-dim);
    opacity:0.45;
    cursor:default;
    display:inline-flex;
    align-items:center;
    gap:7px;
  }
  .navlinks .navlink-disabled::after{
    content:"";
    width:5px; height:5px; border-radius:50%;
    background:var(--bio); opacity:0.8;
  }

  /* SERVICES PAGE */
  .services-hero{ padding-top:190px; padding-bottom:70px; }
  .services-hero .head{ max-width:720px; margin-bottom:0; }

  .services-grid{
    display:grid; grid-template-columns:repeat(4,1fr); gap:1px;
    background:var(--line); border:1px solid var(--line); border-radius:6px; overflow:hidden;
  }
  .service-card{ background:var(--ink-2); padding:32px 28px; transition:background .25s; }
  .service-card:hover{ background:var(--ink-3); }
  a.service-card:hover h3{ color:var(--bio); }
  .service-card .num{ font-family:'IBM Plex Mono', monospace; font-size:0.7rem; color:var(--amber); letter-spacing:0.08em; margin-bottom:14px; display:block; }
  .service-card h3{ font-size:1.05rem; margin-bottom:8px; line-height:1.25; }
  .service-card .taxon{ font-size:0.82rem; display:block; margin-bottom:12px; }
  .service-card p{ color:var(--paper-dim); font-size:0.87rem; margin:0; }

  @media (max-width:1080px){
    .services-grid{ grid-template-columns:repeat(2,1fr); }
  }
  @media (max-width:560px){
    .services-grid{ grid-template-columns:1fr; }
  }

  /* ARTICLE / ENTORNO DETAIL PAGES */
  .breadcrumb{
    font-family:'IBM Plex Mono', monospace; font-size:0.78rem; color:var(--paper-dim);
    text-decoration:none; display:inline-flex; align-items:center; gap:8px; margin-bottom:22px;
  }
  .breadcrumb:hover{ color:var(--bio); }
  .article{ max-width:760px; }
  .article .intro{ font-size:1.05rem; color:var(--paper-dim); max-width:68ch; margin:0 0 16px; }
  .article .intro:last-of-type{ margin-bottom:0; }
  .article a.btn{ margin-top:8px; }
  .article-body{ max-width:760px; }
  .article-block{ padding:44px 0; border-bottom:1px solid var(--line); }
  .article-block:first-child{ padding-top:0; }
  .article-block:last-child{ border-bottom:none; }
  .article-block .num{ font-family:'IBM Plex Mono', monospace; font-size:0.72rem; color:var(--amber); letter-spacing:0.08em; margin-bottom:12px; display:block; }
  .article-block h3{ font-size:1.3rem; margin-bottom:18px; }
  .article-block h4{ font-family:'Space Grotesk', sans-serif; font-size:1rem; color:var(--bio); margin:24px 0 8px; font-weight:600; }
  .article-block h4:first-of-type{ margin-top:0; }
  .article-block p{ color:var(--paper-dim); font-size:0.96rem; margin:0 0 14px; max-width:68ch; }
  .article-block p:last-child{ margin-bottom:0; }
  .article-block ul{ margin:0 0 14px; padding-left:20px; color:var(--paper-dim); font-size:0.96rem; }
  .article-block ul li{ margin-bottom:6px; }
  .article-block strong{ color:var(--paper); font-weight:600; }
  .scenario-grid{ display:grid; grid-template-columns:repeat(2,1fr); gap:1px; background:var(--line); border:1px solid var(--line); border-radius:6px; overflow:hidden; margin:20px 0 22px; }
  .scenario-card{ background:var(--ink-2); padding:18px 20px; }
  .scenario-card .code{ font-family:'IBM Plex Mono', monospace; font-size:0.74rem; color:var(--amber); display:block; margin-bottom:6px; }
  .scenario-card p{ margin:0; font-size:0.88rem; }
  .note-box{
    border:1px dashed var(--line-strong); border-radius:6px; padding:18px 20px; margin-top:8px;
    font-size:0.88rem; color:var(--paper-dim);
  }
  .note-box b{ color:var(--amber); font-weight:500; }
  .source-list{ list-style:none; margin:0; padding:0; }
  .source-list li{ padding:12px 0; border-bottom:1px solid var(--line); font-size:0.88rem; }
  .source-list li:last-child{ border-bottom:none; }
  .source-list a{ color:var(--bio); text-decoration:none; }
  .source-list a:hover{ text-decoration:underline; }
  .source-list span{ display:block; color:var(--paper-dim); margin-bottom:4px; }

  .date-list{ list-style:none; margin:0; padding:0; }
  .date-list li{
    display:flex; justify-content:space-between; align-items:center; gap:20px; flex-wrap:wrap;
    padding:16px 0; border-bottom:1px solid var(--line); font-size:0.95rem;
  }
  .date-list li:last-child{ border-bottom:none; }
  .date-list .date-title{ color:var(--paper); }
  .date-list a{
    color:var(--bio); text-decoration:none; font-family:'IBM Plex Mono', monospace;
    font-size:0.85rem; white-space:nowrap;
  }
  .date-list a:hover{ text-decoration:underline; }

  .data-table{ width:100%; border-collapse:collapse; font-size:0.86rem; margin:8px 0 20px; }
  .data-table th, .data-table td{ padding:10px 14px; border:1px solid var(--line); text-align:left; }
  .data-table th{
    font-family:'IBM Plex Mono', monospace; font-size:0.7rem; text-transform:uppercase;
    letter-spacing:0.06em; color:var(--amber); background:var(--ink-2);
  }
  .data-table td{ color:var(--paper-dim); }
  .data-table-wrap{ overflow-x:auto; }

  @media (max-width:640px){
    .scenario-grid{ grid-template-columns:1fr; }
  }

  /* CONTACT FORM */
  .contact-form{
    border:1px solid var(--line); border-radius:6px; padding:32px;
    background:var(--ink-2);
  }
  .form-row{ margin-bottom:20px; }
  .form-row label{
    display:block; font-family:'IBM Plex Mono', monospace; font-size:0.76rem;
    text-transform:uppercase; letter-spacing:0.06em; color:var(--paper-dim); margin-bottom:8px;
  }
  .form-row input, .form-row textarea{
    width:100%; background:var(--ink); border:1px solid var(--line-strong); border-radius:4px;
    padding:12px 14px; color:var(--paper); font-family:'Inter', sans-serif; font-size:0.95rem;
  }
  .form-row input:focus, .form-row textarea:focus{
    outline:2px solid var(--bio); outline-offset:1px; border-color:var(--bio);
  }
  .form-row textarea{ min-height:150px; resize:vertical; }
  .form-honeypot{ position:absolute; left:-9999px; width:1px; height:1px; overflow:hidden; }
  .form-banner{
    border-radius:6px; padding:16px 20px; margin-bottom:24px; font-size:0.9rem;
  }
  .form-banner.success{ background:rgba(95,225,176,0.1); border:1px solid var(--bio-dim); color:var(--bio); }
  .form-banner.error{ background:rgba(193,87,58,0.12); border:1px solid var(--rust); color:var(--rust); }
  .form-banner a{ color:inherit; text-decoration:underline; }
  @media (max-width:880px){
    .contact-grid[style]{ align-items:stretch !important; }
  }